The year is 2307 A.D. Although fossil fuels have been depleted, humanity has obtained a new source of energy to replace them, in the form of a large-scale solar power generation system based on three huge orbital elevators. However, the benefits of this system are available only to a handful of major powers and their allies. In this world of never-ending conflict, a private armed organization appears, dedicated to the elimination of war through armed force. Its name is Celestial Being, and it is in possession of "Gundam" mobile suits.
At a military testing ground near its orbital elevator, the AEU is conducting a demonstration of its new Enact mobile suit. Among the audience are ace pilot Graham Aker and engineer Billy Katagiri of the Union forces.
As the world reacts with confusion to Celestial Being's announcement, the Gundam Meisters prepare for a second mission which will clarify their intentions. While Allelujah and Tieria descend from space, Setsuna and Lockon make contact with the Celestial Being agent Wang Liu-Mei, and all four Gundam Meisters then join forces for an armed intervention on the island of Ceylon.
The Union ace pilot Graham Aker confronts Setsuna as he leaves Ceylon. Graham's Union Flag mobile suit proves to be no match for Setsuna's Gundam, but he returns from battle to learn that he and Billy Katagiri have been reassigned to a new anti-Gundam investigative taskforce.
The Human Reform League holds a memorial service for the soldiers whose lives were lost in the recent interventions, and reaffirms its opposition to Celestial Being. Sergei Smirnov is placed in command of a new team whose mission is to eliminate the Gundams.
Princess Marina leaves Azadistan and begins a worldwide diplomatic tour to seek help for her impoverished nation. Meanwhile, Kinue Crossroad's younger brother Saji travels into space via the Human Reform League's orbital elevator for an educational study tour.
Celestial Being's activities have reduced the level of conflict in the world, but there are some who do not welcome this change. Among them is the southern European nation of Moralia, whose economy is based on the arms industry and on private military companies (PMCs).
During Celestial Being's intervention in Moralia, Setsuna has been reunited with a figure from his past. His opponent is the mercenary soldier Ali al-Saachez, the same man who once trained Setsuna as a child soldier. Recklessly, Setsuna opens his Gundam's cockpit to confirm his enemy's identity.
An international terror network has begun a campaign of bombing attacks in order to force Celestial Being to halt its armed interventions. While Wang Liu-Mei and Celestial Being's other agents investigate this network, the Ptolemaios crew watch anxiously from the sidelines, and the Gundam Meisters are placed on standby to respond to new attacks.
Four months have now passed since Celestial Being began its armed interventions. The initial chaos has subsided, and life goes on. Louise Halevy's mother visits Tokyo to urge her daughter to come home to Spain, Kinue Crossroad continues her investigation of the mysterious Aeolia Schenberg, and United Nations ambassador Alejandro Corner promises to aid Azadistan by connecting it to the solar power network
In Earth orbit, Sergei's mobile suit team continues its attack on the Ptolemaios and the Gundam Meisters. While Setsuna and Lockon defend their mothership, Allelujah and Tieria find themselves caught in the enemy's trap.
In the aftermath of the orbital battle, the Human Reform League's scientists realize that one of the Gundam Meisters is actually a former test subject of their own Superhuman Institute. Allelujah, meanwhile, has learned that the institute is still in operation.
Although Princess Marina has secured assistance from the United Nations for her troubled homeland, the Kingdom of Azadistan is still in turmoil. A dispute over the introduction of solar power has brought the country's conservative and reformist factions to the brink of civil war.
Setsuna's actions have prevented a coup d'etat in the Kingdom of Azadistan, but the conservative leader Massoud Rachmadi remains missing, and the situation continues to worsen. In order to find out who is behind the crisis, Setsuna visits the site of the solar antenna facility, where he encounters the Union ace pilot Graham Aker.
The three superpower blocs begin planning a massive joint military exercise which will serve as a challenge to Celestial Being. Although they recognize this as an obvious trap, the Gundam Meisters have no choice but to respond with an armed intervention, and the Ptolemaios crew descend to Earth to supervise this dangerous mission.
The Union, Human Reform League, and AEU begin their joint military exercise in the Taklamakan Desert. A nearby uranium storage facility has been left undefended as bait for terrorists, and when Celestial Being intervenes to defend the facility, the combined armies of the three superpower blocs attack the Gundams with overwhelming force.
On the brink of defeat, the Gundam Meisters have been rescued by the intervention of Celestial Being's second team. Their rescuers identify themselves as Johann, Michael, and Nena Trinity, the Meisters of the mysterious Gundam Thrones.
The Ptolemaios crew return to space for a rendezvous with Team Trinity. When they finally meet the new Gundam Meisters face to face, Setsuna and his comrades are disturbed by their attitude and behavior.
Team Trinity continues its campaign of armed intervention. Unlike Setsuna and his comrades, they are targeting military facilities of all kinds and exterminating everyone on the scene, even civilians.
Having determined that Team Trinity itself is promoting conflict, Setsuna confronts the Gundam Thrones. Tieria comes to his assistance, and the two Gundam Meisters fight as a team for the first time.
A mysterious benefactor has provided the three superpower blocs with a new weapon against the Gundams. Quickly establishing a military alliance, the superpowers organize an elite fighting force under the aegis of the United Nations.
The United Nations forces have deployed their new weapon, a mobile suit model known as the GN-X which is powered by a pseudo solar reactor. While Sergei Smirnov and Soma Peries lead a GN-X team in pursuit of the Gundam Thrones, another force made up of Union and AEU pilots attacks the Ptolemaios in space.
The United Nations reveals the existence of the GN-X to the world, and announces the launch of a Gundam elimination operation with the code name "Fallen Angels." The operation begins with the renewed pursuit of Team Trinity, who have been forced to abandon their supply base and can now only flee before the enemy's superior numbers.
Alejandro Corner and Ribbons Almark have forced their way into the heart of Veda and taken control of Celestial Being's master computer. But their actions have triggered an unexpected programmed response.
Despite the loss of Lockon, the Ptolemaios crew are united in their determination to keep fighting the United Nations forces. The enemy begins another assault, this time spearheaded by the Alvatore, a giant mobile armor operated by the traitor Alejandro Corner.
The Celestial Being mothership Ptolemaios has been destroyed, and most of the Gundams are damaged beyond repair. Yet Setsuna continues his desperate fight against the overwhelming power of Alejandro Corner's mobile armor and the final weapon hidden within it. The Celestial Being mothership Ptolemaios has been destroyed, and most of the Gundams are damaged beyond repair. Yet Setsuna continues his desperate fight against the overwhelming power of Alejandro Corner's mobile armor and the final weapon hidden within it.
The year is 2312 A.D, five years after the first appearance of Celestial Being, and four years after the organization was destroyed by the United Nations forces. Thanks to the Gundam Meisters, the nations of the world have been united under the Earth Sphere Federation.
With the reappearance of Celestial Being, the A-LAWS begin gathering personnel who have experience fighting the Gundams. Despite their misgivings about the A-LAWS, Kati Mannequin and Soma Peries accept their new positions, promising to tell their friend Sergei Smirnov what they learn about this mysterious organization.
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The A-LAWS prepare a new trap for the reborn Celestial Being at a detention center for anti-government elements. This detention center holds the former Gundam Meister Allelujah Haptism, and the A-LAWS anticipate that Celestial Being will attempt to rescue their former comrade.
Setsuna and his comrades have successfully overpowered the A-LAWS and freed Allelujah and Marina from the Federation's detention center. When Marina asks them to return her to Azadistan, they set a course back to the Middle East.
Having been rescued from near defeat by the Katharon resistance group, the Ptolemaios 2 now takes refuge at a Katharon base in the Arabian desert. Resistance leader Klaus Grad offers to join forces with Celestial Being against their common enemy, but the crew are reluctant to ally themselves with Katharon.
The Katharon base has been devastated by the A-LAWS attack. The words of a dying soldier remind Sumeragi of a traumatic incident from her past, and she passes out.
Sumeragi has regained consciousness, and thanks to her tactical plan the Ptolemaios 2 is able to escape the battlefield. However, Allelujah's Gundam has been shot down along with Soma Peries, and his comrades cannot locate him.
Tieria is shocked at the sudden appearance of his lookalike Regene Regetta. He is further disturbed by Regene's revelations about the Innovators.
Setsuna and Tieria are confronted by Ali al-Saachez and his new Arche Gundam. A furious Tieria tries to avenge Lockon, but is overwhelmed by the enemy's fighting strength. From Tieria's behavior, Lyle learns that al-Saachez is the man who killed his brother.
The Federation's plan to forcibly reorganize the Middle East is opposed by the kingdom of Suille, the strongest of the Middle Eastern nations. Concerned by this resistance, the Federation decides to dispatch the A-LAWS to Suille, and Sergei Smirnov is ordered to observe the situation from the country's borders.
While Ian Vashti and his fellow Celestial Being mechanics prepare the O Raiser, a support machine for Setsuna's 00 Gundam, the A-LAWS deploy a devastating new weapon of their own. Tieria is outraged by the actions of Ribbons Almark and the A-LAWS, and he finally reveals the existence of the Innovators to the crew of the Ptolemaios 2.
The Ptolemaios 2 attempts to make a hasty escape from the attacking A-LAWS. But the Gundam Meisters are hard-pressed by Kati Mannequin's tactics and the enemy's new mobile suits. In the midst of this battle, the 00 Raiser's Twin Drive reveals its true power.
The Katharon space fleet has begun an assault on the satellite weapon Memento Mori, and is now under attack by A-LAWS mobile suits. The Gundam Meisters join the fray, and Setsuna engages the Innovator Hiling Care. But is this just a trick to lure Setsuna away?
The Ptolemaios 2, damaged by the Innovators' attack, descends to Earth. Now separated from his comrades, Setsuna searches for the Ptolemaios 2. But instead, he finds Ali al-Saachez and his Arche Gundam waiting for him.
The Ptolemaios 2 has received supplies from Katharon, and repairs are now in progress. But it cannot break through the Federation's siege without the fighting strength of the 00, and Setsuna has not yet returned to the ship. While the Ptolemaios 2 wages a defensive battle, news arrives of a coup d'etat within the Federation itself.
The A-LAWS have abruptly retreated after receiving word of a coup d'etat within the Federation. Hoping to reunite with his comrades, Setsuna sets the 00 on a course for Africa. But waiting for him there is the mysterious Mister Bushido, who has reappeared with a brand new mobile suit.
A crisis is unfolding, but the Ptolemaios 2, which has not yet been fully repaired, is unable to break the deadlock. The wounded Setsuna returns to battle despite his injuries, asking Saji to help him save countless innocent lives.
Four months have passed since the "Break Pillar" incident, the near-collapse of an orbital elevator. The A-LAWS leaders receive word that Celestial Being has resumed activity after months of silence.
As Saji Crossroad searches for Louise Halevy amid the chaos of battle, Soma Peries hunts for Andrei Smirnov in order to avenge Sergei's death. Setsuna confronts the Innovator Hiling Care and unleashes the power of the Twin Drive. The particles released by the 00 Raiser spread out across the entire battlefield.
The Innovator Revive Revival is now a prisoner aboard the Ptolemaios 2, but he sneers at the questions of its crew, and gives them nothing but evasive replies. Suddenly, the crew receive word that crewmember Mileina Vashti has been taken hostage.
The Ptolemaios 2 has received a message containing coordinates for a point at Lagrange 5. The crew are suspicious, but Setsuna suggests they set course for this mysterious location. Ian Vashti agrees, since this will enable them to collect new equipment from Celestial Being's nearby secret factory.
The A-LAWS and Federation Forces are gathering a huge fleet to exterminate Celestial Being, and their leaders anticipate a final decisive battle. Aboard the Ptolemaios 2, Setsuna and his comrades gather their resolve for the coming battle. They each give voice to their determination, and crewmember Feldt Grace gives Setsuna a flower which once grew in his Middle Eastern homeland.
The Innovator Regene Regetta finally rebels against Ribbons Almark. Meanwhile, Katharon and the Federation Forces have also been drawn into the savage battle between the A-LAWS and Celestial Being. Suddenly, a giant burst of light envelops the battlefield.
As Tieria enters the enemy ship and learns shocking truths, the Ptolemaios 2 fights a defensive battle against the unrelenting enemy assault. One by one, our heroes fall. Setsuna cries out as if responding to the fading light of their lives.
Setsuna begins a final decisive duel. Lyle's intentions, Allelujah's hopes, and Tieria's thoughts flow through the darkness of space. The particles from the 00 Raiser envelop the area around them...